“…Several hypotheses have been formulated for the development of bottom anoxia in the Eastern Mediterranean Sea, mainly linked to higher productivity levels and the weakening/interruption of deep water formation (Rohling, 1994;Cramp and O'Sullivan, 1999). In any case, the exceptional African Monsoon activity, and the consequent strong Nile (and other presently buried North African) river runoff, was a prime forcing factor of this phenomenon (Rossignol-Strick, 1983, 1985Hilgen, 1991;Lourens et al, 1996;Rohling et al, 2002aRohling et al, , 2004Marino et al, 2007Marino et al, , 2009). Thus, the two peaks of G. quadrilobatus may be an indirect response to the tropical climate influence.…”