“…However, similar endeavors to study mobile machines in athermal fluids such as nonequilibrium granular materials have been initiated only recently [1], mainly because the nonequilibrium features make developing a governing equation of the system a challenging task [2]. Among the finite attempts tackling this challenge in the literature, except some artificial designs [3][4][5][6][7][8], most of them are bio-inspired, such as mimicking sandfish, lizard and snake [4,[9][10][11][12][13][14][15], insects [16] and clam that uses an intricate way to move forward by swallowing and discharging sands [17,18]. Learning from the nature has the benefit of always having an original counterpart that has been optimised through long evolution to compare with, and the mimicked designs are guaranteed to function under known conditions.…”