“…The genus Periophthalmus is the richest mudskipper genus (Jaafar et al ., ). It is basal to all oxudercine and amblyopine genera so far examined (Thacker, ; Agorreta & Rüber, ; Agorreta et al ., ; Tornabene et al ., ) and includes cryptic species with conserved morphologies, suggestive of a long evolutionary history (>30 M years) in coastal wetlands (Polgar et al ., ). Periophthalmus has the widest geographic distribution, along the African Atlantic Ocean coasts eastward to the Indo‐West‐Pacific region (Polgar et al ., ) and its species live in a range of intertidal and supratidal habitats, from unvegetated mud banks and open tidal mudflats (Baeck et al ., ; Polgar & Crosa, ; Polgar & Bartolino, ; Takita et al ., ) to mangrove forests and freshwater swamps (Larson, ; Polgar et al ., ).…”