“…During recent decades, pools and fluxes of nitrogen (N), phosphorus (P) and carbon (C), in UK ecosystems have been transformed (Galloway et al, 2008;Vitousek et al, 1997) by the spread and fertiliser-based intensification of agriculture (Jenkinson, 2001), by atmospheric pollution (Fowler et al, 2005;Goulding et al, 1998), and now potentially by climate change (Stuart et al, 2011;Watts et al, 2015). Many terrestrial ecosystems that were previously nearly-closed with respect to N and P now permit considerable export of these nutrients (Galloway et al, 2008;Smil, 2000;Withers et al, 2001), leading to eutrophication and acidification of surface waters (Bennett et al, 2001).…”