“…Gradient analysis is a way to compare quantitatively the composition and abundance of taxa among localities, and, in the marine subfossil and fossil record, is a valuable tool for characterizing the spatio-temporal distribution of organisms according to paleoenvironmental variables (e.g., Cisne and Rabe, 1978;Rabe and Cisne, 1980;Cisne et al, 1982;Springer and Bambach, 1985;Anstey et al, 1987;Miller, 1988;Anstey and Rabbio, 1990;Miller et al, 1992;Lafferty et al, 1994;Patzkowsky, 1995;Dattilo, 1996). A method of high-resolution faunal gradient analysis applied by Holland et al (2001) and Miller et al (2001) and modified by Webber (2002Webber ( , 2004 has provided a valuable means of characterizing the signal of biological variation for strata in the Kope and lower Fairview formations of the type Cincinnatian Series (Upper Ordovician) at a regional scale (Fig.…”