JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact support@jstor.org.. However, the excavating guild includes twice as many species in Guatemala as in either of the northern study sites. Two of the three "additional' species in Guatemala use a configuration of foraging methods and perches not used by northern woodpeckers. Hence the larger number of tropical woodpecker species can be attributed partly to the greater range of resources available in the structurally complex rain forest.