“…In another study, a sapling species richness of only 21 % was found in gaps smaller than 50 m 2 (Hubbell et al, 1999). In these studies, the main causes of species diversity in gap regeneration probably were: i) less adult tree species at the gap edge of small gaps, or/and, ii) an early high degree of speciesselective self-thinning, due to intra and interspecific competition-induced mortality in small gaps, during the course of population development in tree community (Lepš & Kindlmann, 1987;Getzin, Wiegand, Wiegand, & He, 2008;García-Domingo & Saldana, 2013;Wehenkel, Brazão-Protázio, Carillo-Parra, Martínez-Guerrero, & Crecente-Campo, 2015), because of different seed production rate, dispersal, and vigour and shade tolerance of seedlings and saplings of different tree species (Canham, 1989). In contrast, the competition-induced mortality in the larger gaps was probably lower due to lower over story competition on understory gap regeneration.…”