A New Subfamily of Aulacigastridae (Diptera: Schizophora), with a Discussion of Aulacigastrid Classification

Hennig’s orphans revisited: Testing morphological hypotheses in the “Opomyzoidea” (Diptera: Schizophora)

“…This classification partially follows results of previous studies (e.g. McAlpine, 1978McAlpine, , 1983, but other proposed groupings were not explicitly justified. D. K.…”

“…Later, D.K. McAlpine (1978McAlpine ( , 1983) transferred all three genera to the Periscelididae on the basis of similar antennal structure, a placement that has been followed by recent authors (Grimaldi and Mathis, 1993;Baptista and Mathis, 1994;Mathis and Papp, 1998;Mathis, in press). …”

“…1) that are grouped into four suprafamilies: Clusioinea (Clusiidae and Acartophthalmidae), Agromyzoinea (Agromyzidae, Fergusoninidae, and Odiniidae), Opomyzoinea (Opomyzidae and Anthomyzidae), and Asteioinea (Asteiidae, Aulacigastridae, Xenasteiidae, Teratomyzidae, Neurochaetidae, and Periscelididae). Two families named more recently have been associated with Opomyzoidea after McAlpine (1989): Neminidae (Freidberg, 1994; described as a subfamily of Aulacigastridae by McAlpine, 1983) and Marginidae (newly described and provisionally placed in Opomyzoidea by McAlpine, 1991). In addition, McAlpine (1989;following Hennig, 1971) suggested that Paraleucopis Malloch (currently in Chamaemyiidae) and the unplaced Neotropical genera Mallochianamyia Santos-Neto (= Gayomyia Malloch; see Wheeler, 2000) and Schizostomyia Malloch probably represent a separate lineage closely related to Asteiidae.…”

“…For example, the diminutive Xenasteiidae were independently described from Mediterranean and Pacific seashores less than three decades ago (Hardy, 1980;Papp, 1980) and are still largely unknown. Neminidae, dubbed ''nobody flies" by their discoverer (Latin nemo = ''nobody"; McAlpine, 1983McAlpine, , 1998, are now known from southern Africa, Australia, and New Guinea, but have been very rarely collected, and nothing is known about their biology. Although easily recognized by the laterally elongate head produced into eyestalks (Mathis and Rung, 2004), the bizarre periscelidid genus Diopsosoma, with only one included species, D. primum Malloch, is still known only from the type series collected in Peru in 1931 (Mathis and Rung, 2004).…”

A NEW SPECIES AND SUBGENUS OF PERISCELIS LOEW FROM AUSTRALIA (DIPTERA: PERISCELIDIDAE)

“…In this paper, McAlpine's (1 978, 1983) expanded concept of the family is adopted. McAlpine (1978McAlpine ( , 1983) included a few genera such as Cyamops, Planinasus and Stenomicra that were previously assigned to the family Aulacigastridae. These genera are closely related and are here recognised as the subfamily Stenomicrinae.…”