There are many people without whom I could not have successfully completed my thesis, and I am eternally grateful for all of their contributions. Firstly, I must thank my advisor, Dr. Jill Pruetz, for her endless advice and support. Her patience and understanding throughout the many stages of my project were invaluable, and I continue to be inspired by her experience and passion for primatology. My committee members, Dr. Max Viatori and Dr. Doug Gentile provided innovative insight and critiques in developing and refining my thesis, which I truly appreciate. I also thank the Department of Anthropology at Iowa State for their financial support and honoring me with the Nancy R. Coinman Award for anthropological research, allowing me to contribute to this incredible project. The directors of the Ape Cognition and Conservation Initiative, Drs. Jared Taglialatela and William Hopkins, provided priceless guidance from their many impressive years of research experience. Their commitment to producing reliable scientific research, as well as their dedication to the five incredible apes at ACCI, continually enriched this study. I am indebted to Dr. Ken Schweller for his endless ideas, help, and patience in solving all of ACCI's programming and computer needs, especially through developing specialized training and testing protocols for the bonobos. There would not even be a completed project without the staff members of ACCI who were present during my study. I thank you for your daily help working with the bonobos, shifting them between rooms, and keeping track of Kanzi's grape calories for many months of data collection. I know the apes all appreciate your care even more than I do. I also thank Eric Mittman, of the Iowa State Statistics department for his assistance and patience in analyzing my data. I am