“…We test these hypotheses in the custard apple family (Annonaceae, Magnoliales), a major vertebrate‐dispersed, globally distributed plant family, typically confined to the rain forest biome (Keßler, ). Annonaceae comprises c. 2,400 species and has been well‐studied from phylogenetic (Chatrou et al., ; Guo, Tang, Thomas, Couvreur, & Saunders, ), evolutionary (Couvreur et al., ; Pirie, Maas, Wilschut, Melchers‐Sharrott, & Chatrou, ) and biogeographical (Couvreur et al., ; Doyle, Sauquet, Scharaschkin, & Le Thomas, ; Thomas et al., ; Zhou, Su, Thomas, & Saunders, ) points of view. Furthermore, almost all Annonaceae species are dispersed by animals, ranging from birds, bats, primates, small‐bodied mammals (e.g., rodents), and, in some cases, fish (Keßler, ).…”