“…Such a pattern, already documented in red deer (Cervus elaphus; Drucker et al, 2003;Drucker et al, 2008), is consistent with the changes in the forest cover as a result of vegetation succession in Europe in the Early Holocene, but it is not compatible with the reduction of the forest cover as a result of agricultural activities following the Neolithic and subsequent periods (Woodbridge, Fyfe, Roberts, Mazier, & Davis, 2018). Thus, aurochs and European bison, preadapted to open or mixed habitats Hall, 2008), became classical refugee species sensu Kerley et al (2012) after that were introduced mainly to forests . This has continued through the last 2,000 years bp, when the scale of forest reduction was the largest (Kaplan et al, 2009).…”