“…For example, studies on the scaling of population fluctuations revealed the central role of temporal environmental variability in shaping ecological communities (Chisholm et al, 2014;Jabot & Lohier, 2016;Kalyuzhny, Schreiber, et al, 2014;Kalyuzhny, Kadmon, & Shnerb, 2015), and studies focusing on long-term changes in abundance and diversity revealed that population-level regulation is often weak (Kalyuzhny, Seri, et al, 2014;Knape & de Valpine, 2012;Ziebarth, Abbott, & Ives, 2010), while total-abundance and species diversity are indeed regulated (Brown, Ernest, Parody, & Haskell, 2001;Goheen, White, Ernest, & Brown, 2005;Gotelli et al, 2017;Magurran & Henderson, 2018). Moreover, in recent years there is an increasing interest in understanding richness trends and compositional turnover, partly motivated by concerns over the effect of anthropogenic activities on ecological communities (Elahi et al, 2015;Magurran et al, 2018;McGill et al, 2015;Vellend et al, 2013). Several studies have shown that while some local communities show richness trends, negative and positive changes may cancel out in multiple communities worldwide (Dornelas et al, 2014;Vellend et al, 2013).…”