“…The LMMCs serve several important functions. Along with glycoproteins, they confer hygroscopicity, causing the droplets' size and performance to change over the course of a day as they track environmental humidity (Figure d; Jain et al, ; Opell, Clouse, & Andrews, ; Opell, Jain, et al, ), they maintain glycoprotein structure and solvate glycoprotein, enhancing its surface interaction (Sahni et al, ), and they remove interfacial water from a droplet's contact footprint, enhancing adhesion (Singla, Amarpuri, Dhopatkar, Blackledge, & Dhinojwala, ). Optimal adhesion of an araneoid glue droplet is achieved when the viscosity of the droplet's glycoprotein is low enough to spread on a surface to establish sufficient adhesive contact, but high enough to ensure that the glycoprotein will cohere as it extends, thereby transferring adhesive force to the thread's axial lines (Figure c; Amarpuri, Zhang, Blackledge, & Dhinojwala, ; Amarpuri, Zhang, et al, ).…”