“…Coordinated embryo, endosperm, and maternal seed coat development is necessary for normal seed formation. However, seed abortion, a typical postzygotic reproductive barrier preventing seed development or seed germination, is widely observed in interspecific and interploidy crosses (Baek et al, 2016;Coughlan et al, 2020;Oneal et al, 2016;Rebernig et al, 2015;Roth et al, 2018;Scott et al, 1998;Sekine et al, 2013;Tonosaki et al, 2018;). Studies on Arabidopsis, Solanum, Mimulus, and other species have strongly suggested that endosperm developmental failure is the primary cause of seed abortion, as this failure leads to embryo arrest and seed abortion (Hehenberger et al, 2012;Rebernig et al, 2015;Scott et al, 1998;Sekine et al, 2013;Tonosaki et al, 2018;Oneal et al, 2016;Coughlan et al, 2020;Roth et al, 2018).…”