Abstract. High resolution, downscaled climate model data are used in a wide variety of applications in environmental sciences. Here we present the CHELSA-TraCE21k downscaling algorithm to create global monthly climatologies for temperature and precipitation at 30 arcsec spatial resolution in 100 year time steps for the last 21,000 years. Paleo orography at high spatial resolution and at each timestep is created by combining high resolution information on glacial cover from current and Last Glacial Maximum (LGM) glacier databases with the interpolation of a dynamic ice sheet model (ICE6G) and a coupling to mean annual temperatures from CCSM3-TraCE21k. Based on the reconstructed paleo orography, mean annual temperature and precipitation was downscaled using the CHELSA V1.2 algorithm. The data were validated by comparisons with the glacial extent of the Laurentide ice shield based on expert delineations, proxy data from Greenland ice cores, historical climate data from meteorological stations, and a dynamic simulation of species a distribution throughout the Holocene. Validations show that CHELSA TraCE21k output creates a reasonable representation of the distribution of temperature and precipitation through time at a high spatial resolution, and simulations based on the data are capable of detecting effective LGM refugia of species.

Aims: Ellenberg-type indicator values are expert-based rankings of plant species according to their ecological optima on main environmental gradients. Here we extend the indicator-value system proposed by Heinz Ellenberg and co-authors for Central Europe by incorporating other systems of Ellenberg-type indicator values (i.e., those using scales compatible with Ellenberg values) developed for other European regions. Our aim is to create a harmonized data set of Ellenberg-type indicator values applicable at the European scale.Methods: We collected European data sets of indicator values for vascular plants and selected 13 data sets that used the nine-, ten-or twelve-degree scales defined by Ellenberg for light, temperature, moisture, reaction, nutrients and salinity. We compared these values with the original Ellenberg values and used those that showed consistent trends in regression slope and coefficient of determination. We calculated the average value for each combination of species and indicator values from these data sets. Based on species' co-occurrences in European vegetation plots, we also calculated new values for species that were not assigned an indicator value. Results: We provide a new data set of Ellenberg-type indicator values for 8908European vascular plant species (8168 for light, 7400 for temperature, 8030 for

Abstract. High-resolution, downscaled climate model data are used in a wide variety of applications across environmental sciences. Here we introduce a new, high-resolution dataset, CHELSA-TraCE21k. It is obtained by downscaling TraCE-21k data, using the “Climatologies at high resolution for the earth's land surface areas” (CHELSA) V1.2 algorithm with the objective to create global monthly climatologies for temperature and precipitation at 30 arcsec spatial resolution in 100-year time steps for the last 21 000 years. Paleo-orography at high spatial resolution and for each time step is created by combining high-resolution information on glacial cover from current and Last Glacial Maximum (LGM) glacier databases and interpolations using data from a global model of glacial isostasy (ICE-6G_C) and a coupling to mean annual temperatures from TraCE21k (Transient Climate Evolution of the last 21 000 years) based on the Community Climate System Model version 3 (CCSM3). Based on the reconstructed paleo-orography, mean annual temperature and precipitation were downscaled using the CHELSA V1.2 algorithm. The data were validated by comparisons with the glacial extent of the Laurentide ice sheet based on expert delineations, proxy data from Greenland ice cores, historical climate data from meteorological stations, and a dynamic simulation of species distributions throughout the Holocene. Validations show that the CHELSA-TraCE21k V1.0 dataset reasonably represents the distribution of temperature and precipitation through time at an unprecedented 1 km spatial resolution, and simulations based on the data are capable of detecting known LGM refugia of species.

Bryophytes are a diverse group of organisms with unique properties, yet they are severely underrepresented in plant trait databases. Building on the recently published European Red List of bryophytes and previous trait compilations, we present the Bryophytes of Europe Traits (BET) data set, including biological traits such as those related to life history, growth habit, sexual and vegetative reproduction; ecological traits such as indicator values, substrate and habitat; and bioclimatic variables based on the species' European range. The data set includes values for 65 traits and 25 bioclimatic variables, containing more than 135,000 trait values with a completeness of 82.7% on average. The data set will enable future studies in bryophyte biology, ecology and conservation, and may help to answer fundamental questions in bryology.

Aims: To develop a consistent ecological indicator value system for Europe for five of the main plant niche dimensions: soil moisture (M), soil nitrogen (N), soil reaction (R), light (L) and temperature (T). Study area: Europe (and closely adjacent regions). Methods: We identified 31 indicator value systems for vascular plants in Europe that contained assessments on at least one of the five aforementioned niche dimensions. We rescaled the indicator values of each dimension to a continuous scale, in which 0 represents the minimum and 10 the maximum value present in Europe. Taxon names were harmonised to the Euro+Med Plantbase. For each of the five dimensions, we calculated European values for niche position and niche width by combining the values from the individual EIV systems. Using T values as an example, we externally validated our European indicator values against the median of bioclimatic conditions for global occurrence data of the taxa. Results: In total, we derived European indicator values of niche position and niche width for 14,835 taxa (14,714 for M, 13,748 for N, 14,254 for R, 14,054 for L, 14,496 for T). Relating the obtained values for temperature niche position to the bioclimatic data of species yielded a higher correlation than any of the original EIV systems (r = 0.859). The database: The newly developed Ecological Indicator Values for Europe (EIVE) 1.0, together with all source systems, is available in a flexible, harmonised open access database. Conclusions: EIVE is the most comprehensive ecological indicator value system for European vascular plants to date. The uniform interval scales for niche position and niche width provide new possibilities for ecological and macroecological analyses of vegetation patterns. The developed workflow and documentation will facilitate the future release of updated and expanded versions of EIVE, which may for example include the addition of further taxonomic groups, additional niche dimensions, external validation or regionalisation. Abbreviations: EIV = Ecological indicator value; EIVE = Ecological Indicator Values for Europe; EVA = European Vegetation Archive; GBIF = Global Biodiversity Information Facility; i = index for taxa; j = index for EIV systems; L = ecological indicator for light; M = ecological indicator for moisture; N = ecological indicator for nitrogen availability; R = ecological indicator for reaction; T = ecological indicator for temperature.