“…Benthic invertebrates were collected once per year in late summer/early fall by Crawford and others (1992) and then twice a year during subsequent studies (Voelker and Renn, 2000;Voelker, 2004Voelker, , 2012Voelker, , 2014. Samples for the latter studies were collected during periods of relatively low-flow, steady-state streamflow conditions in the spring (May or June), and in the late summer/early fall (September and October) unless streamflow conditions resulted in sampling-schedule changes.…”