“…However, like roofs, they show very extreme environmental conditions, with flora adapted to a hostile environment that includes limited nutrients, water, and soil accumulation due to highly compacted and thin substrates, extreme light and temperature conditions, a variable soil pH, changing water conditions, eutrophication, and disturbances (trampling, intensive management, pets) [88][89][90][91]. Locally, even in this harsh environment, the heterogeneity of abiotic conditions can enhance species diversity [92].…”