“…Early successional cyanobacterial crusts are light in color, with a small part of fine particles, low protective ability to water and wind erosion, poor nutrients and water retention (Housman et al, 2006;Lan et al, 2010a,b). With crust development and succession, dark-colored lichens or mosses later colonize the soil surface (Lange et al, 1992;Zaady et al, 2000;Lan et al, 2010a). Compared with cyanobacterial crusts, later successional lichen or moss crusts have the higher biological metabolic efficiency and protective ability; hence better topsoil microenvironments are expected in the later BSCs (Castenholz and Garcia-Pichel, 2000;Redfield et al, 2002;Housman et al, 2006).…”