New Spanish Late Cretaceous eusuchian reveals the synchronic and sympatric presence of two allodaposuchids

Abstract

The recently described clade Allodaposuchidae includes European eusuchian crocodyliforms restricted to the Late Cretaceous (Campanian and Maastrichtian). A new allodaposuchid crocodyliform is here described based on two specimens from the upper Campanian-lower Maastrichtian fossil site of Lo Hueco (Cuenca, Spain). This new taxon, Agaresuchus fontisensis gen. et sp. nov., is described by two complete skulls and a lower jaw associated with one of them. This new species can be distinguished unambiguously from Lohuecosuchus megadontos, the other allodaposuchid known from the same fossil site. The presence of two allodaposuchid crocodyliforms in Lo Hueco allows the recognition of the synchronic and sympatric existence of two representatives of this clade for the first time. The new genus Agaresuchus, comprises a previously described Iberian allodaposuchid species,-Allodaposuchus‖ subjuniperus, as Agaresuchus subjuniperus, new combination. Romania. A nearly complete skull from Oarda de Jos (Hateg Basin, Romania) was later referred to the species (Delfino et al., 2008), significantly improving the knowledge on this taxon. Several subsequently described crocodyliforms from Campanian and Maastrichtian sites of the Iberian Peninsula and southern France were also identified as allodaposuchids (see Narváez et al., 2015). Most of them correspond to forms described in recent years. The Iberian forms include Allodaposuchus subjuniperus Puértolas

A review of the Paleogene eusuchian crocodyliform Borealosuchus wilsoni (Mook, 1959) from western North America

“…The position of Borealosuchus in the strict consensus tree as sister to the clade containing Planocraniidae and Brevirostres is consistent with placements found by (Brochu, 1997;Brochu, 2004;and Narváez et al, 2016).…”

Borealosuchus (Crocodylia) from the early Campanian Mooreville chalk reveals new insights into the late Cretaceous fauna of Alabama and the origin of Crocodylian lineages

“…Less resolution was recovered within Allodaposuchidae, the sister group relationships between the two species of Agaresuchus and the two species of Lohuecosuchus are not recovered. This may be due to less extensive sampling of basal eusuchians and advanced neosuchians in this analysis than in Narváez et al (2015Narváez et al ( , 2016.…”

“…Arenysuchus was thought to be crocodyloid when initially coded into this matrix but has now been recovered as an allodaposuchid in several analyses (Puértolas et al, 2011;Narváez et al, 2016). Additionally, all of Allodaposuchia has been recovered as sister to Borealosuchus + Planocraniidae within Crocodylia (Blanco et al, 2015), though other analyses consistently recover this group as a basal eusuchian clade (Narváez et al, 2015(Narváez et al, , 2016 Analyses using different matrices that include mostly taxa outside of Crocodylia and characters focused on basal relationships commonly recover topologies with Borealosuchus just outside of Crocodylia (e.g. Pol et al, 2009;Figueiredo et al, 2011).…”

“…More non-crocodylian eusuchians are being to be added to the matrix of Brochu (2013), especially allodaposuchids (Narváez et al, 2015(Narváez et al, , 2016. This more comprehensive taxon sampling may help to increase the consistency in the placement of Borealosuchus.…”

“…Allodaposuchidae is sister to Crocodylia (Narváez et al, 2016;Mateus et al 2018), or a more inclusive Eusuchia (including paralligatorids and atoposaurids) is sister to Crocodylia (Turner and Pritchard, 2015;Turner, 2015 (Narváez et al, 2015). Borealosuchus sp.…”