“…The high biodiversity of southern China is due in large part to the extreme physiographical heterogeneity of its mountain ranges (Fan et al , 2018;Xu et al, 2017;Yang, Dick, Yao, & Huang, 2016). Often oriented in either north-south or northeast-southwest directions (Hou, 1983;Wang, 1992a;Wang, 1992b;Ying, 2001), these topographically diverse ranges have been suggested to serve as either geographical barriers or colonization corridors for various plant species (Gong et al, 2016;Tian et al, 2018;Xiong, Wu, & Zhang, 2019). Their local habitat uniqueness has been attributed to complex topography correlated with longitudinal or steep elevational gradients (Qiu, Zeng, Chen, Zhang, & Zhong, 2013;Wang, Fang, Tang, & Shi, 2012).…”