“…The development of tip dating with more complex mechanistic tree models, such as the fossilized birth–death (FBD) tree [ 65 , 66 ] and its subsequent variations—such as the skyline FBD [ 13 , 14 , 15 , 16 , 17 , 18 , 19 ], which enables speciation, extinction, and fossilization parameters to be changed piecewise across the tree—allowed fossil species to be analysed in conjunction with and within the same analytical framework as extant taxa using Bayesian phylogenetics. This has been particularly useful for palaeontologists, who have revitalizing the use of morphological data to elucidate the dynamics of evolution across the tree of life [ 16 , 17 , 49 , 51 , 52 , 63 , 67 , 68 , 69 ] ( Figure 1 ). These analyses can be carried out with morphological datasets alone, in what is called the morphological clock [ 16 , 17 , 51 , 52 , 63 , 70 ], which adds another method of reconstructing evolution to the palaeontologist’s toolbox.…”