“…DOI: 10.1038/sj/hdy/6800003 species can be studied by analysing pollen (eg, Huntley and Birks, 1983;Gliemeroth, 1995). For animals, sufficient remains only exist for some groups, such as beetles or terrestrial gastropods (eg, Coope, 1970Coope, , 1994Hertelendy et al, 1992); for the great majority of invertebrates, hardly any evidence of this kind is available.…”