“…The varied canopy closure was primarily maintained by natural factors such as fires, grazing of large herbivores and windthrows (Bengtsson, Nilsson, Franc, & Menozzi, ; Bradshaw & Hannon, ; Hultberg, Gaillard, Grundmann, & Lindbladh, ; Vera, ). Later, the natural disturbances were supplemented or gradually replaced by traditional forest management including, for example, pasture grazing, coppicing and burning (Jamrichová et al., ; Kirby & Watkins, ; Rackham, ; Szabó, Müllerová, Suchánková, & Kotačka, ). In the past two centuries, however, the remaining natural disturbance factors have largely been suppressed and the traditional management has been replaced by the practices of modern forestry (Bürgi, ; Brunet, Felton, & Lindbladh, 2012).…”