“…Thirdly, the gravel-dependent changes in soil moisture and temperature may influence soil enzyme activity (Wu et al, 2011), which is an important mediator for organic matter degradation, mineralization and nutrient recycling (Huang et al, 2013;Tripathy et al, 2014). Studies by Corti et al (1998Corti et al ( , 2002 found that rock fragments (skeleton or gravel) in stony soil with alteration contained substantial amount of organic carbon. However, SOC in the rock fragments (skeleton or gravel) was often neglected in the calculation of total soil C and N due to soil nutrition analysis mainly focused on the fine earth fraction (less than 2 mm).…”