2004
DOI: 10.1038/nature02855
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Abstract: Tyrannosauroids are one of the last and the most successful large-bodied predatory dinosaur groups, but their early history remains poorly understood. Here we report a new basal tyrannosauroid from the Early Cretaceous Yixian Formation of western Liaoning, China, which is small and gracile and has relatively long arms with three-fingered hands. The new taxon is the earliest known unquestionable tyrannosauroid found so far. It shows a mosaic of characters, including a derived cranial structure resembling that o… Show more

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Cited by 203 publications
(231 citation statements)
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“…1A). These structures can be divided into two categories: short, slender filamentous feathers that are similar to those reported in many other nonavian theropods (1,(5)(6)(7)(11)(12)(13), and elongated broad filamentous feathers (EBFFs) that are novel. The EBFFs are preserved ventral to the posterior half of the mandible, dorsal to the posterior portion of the skull and the anterior portion of the neck (Fig.…”
mentioning
confidence: 63%
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“…1A). These structures can be divided into two categories: short, slender filamentous feathers that are similar to those reported in many other nonavian theropods (1,(5)(6)(7)(11)(12)(13), and elongated broad filamentous feathers (EBFFs) that are novel. The EBFFs are preserved ventral to the posterior half of the mandible, dorsal to the posterior portion of the skull and the anterior portion of the neck (Fig.…”
mentioning
confidence: 63%
“…They occur only on portions of the head, neck, and tail, body areas that commonly bear integumentary display structures in extant birds (16) and mammals (17). These features suggest that the EBFFs might have been used in display, whereas other types of filamentous feathers in nonavian theropods are more likely to have functioned in thermoregulation (2,11,12). Integumentary display using highly modified pennaceous feathers has been documented in basal avians (18) and even in nonavian maniraptorans (19), but no filamentous feathers have been shown to function in display.…”
Section: Discussionmentioning
confidence: 99%
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“…The origin and evolution of feathers have been of great interest (3,4), particularly with the recent discoveries of feathered dinosaurs in Northern China (5,6). From the many intermediate feather forms, we learned that feathers evolved through stepwise evolutionary novelties to produce diverse morphology with new functions (7,8).…”
mentioning
confidence: 99%
“…The protofeathers of taxa such as Beipiaosaurus, Yutyrannus, and Dilong (as well as several other theropod and orntihischian dinosaurs not from the Jehol Biota [Rauhut et al 2012, Zelenitsky et al 2012, Godefroit et al 2014) are of much more robust dimensions and quill-like morphology than the down of chicks (Xu et al 1999(Xu et al , 2004(Xu et al , 2009a(Xu et al , 2009b(Xu et al , 2009c(Xu et al , 2012. As such, there are no data to support Feduccia's (2013) proposed correlations between the down of chicks and protofeathers.…”
Section: Anatomical and Evolutionary Misinterpretationsmentioning
confidence: 99%