“…The Woodbine Formation in north-central Texas preserves a rare Cenomanian (95-100 Ma) ecosystem from the southwestern corner of the ancient landmass of Appalachia. A rich diversity of vertebrate taxa from the Woodbine Formation have been described to date, including dinosaurs (the hadrosauroid Protohadros, a nodosaurid, at least four theropods, and the enantiornithine bird Flexomornis), neosuchian crocodyliforms, turtles, multituberculate mammals, osteichthyan fish, and chondrichthyan fish (Head, 1998;Jacobs and Winkler, 1998;Lee, 1997aLee, , 1997bMain, 2013;Main et al, 2011;McNulty and Slaughter, 1962;Noto, 2015;Tykoski and Fiorillo, 2010). Historically, the clade-level assignment of taxa preserved within the formation was difficult to assess, largely because fossils, while common, are often found isolated, fragmented, disarticulated, and poorly preserved.…”