“…31,32 In this paper, the heterogeneity of habitat is defined by habitat complexity, which refers to the morphological characteristics within a structure itself or the heterogeneity in the arrangement of objects in space and plays an important role in dynamical consequences of predatorprey systems. 31,[35][36][37][38][39][40][41][42][43][44][45][46] Habitat complexity is found in almost all ecological systems; a case in point is that marine habitat becomes complex in the presence of oyster and coral reefs, mangroves, sea grass beds and salt marshes. 35 In lakes, habitat heterogeneity is most commonly present in the form of littoral zone vegetation or depth-gradient diversity.…”