“…From the species list, a species character matrix was built for 104 species of Lejeuneaceae, with records of the following: presence of dark pigmentation, ability to convolute leaves when dry, possession of thickened cell walls, reproduction mode (monoicous or dioicous), occurrence of asexual reproduction and epiphyllous habit. All traits were scored as present (1) or absent (0) at species level, and reproductive system as monoicous (1) or dioicous (0) (or both, 0-1, in four species), based mainly on taxonomic literature ( Bastos, 2008 ; Bastos, 2017 ; Bernecker-Lücking, 1998 ; Bischler, 1967 ; Bischler, 1969 ; Dauphin, 2003 ; Gradstein, 1994 ; Gradstein, 1997 ; Gradstein & Costa, 2003 ; He, 1999 ; Ilkiu-Borges & Gradstein, 2008 ; Ilkiu-Borges & Lisboa, 2004 ; Jovet-Ast, 1953 ; Piippo, 1986 ; Reiner-Drehwald, 1995 ; Reiner-Drehwald, 2009 ; Reiner-Drehwald & Goda, 2000 ; Schuster, 1980 ). A few cases interpreted as doubtful by the author were scored with an X, and the species was excluded from the analysis of that trait.…”