“…Several studies have focused vertical stratification in Anura (Arroyo et al, 2008;Muñoz-Guerrero, 2007;Guayasamin and Funk, 2009;de Menezes Gondim, 2013;Fonseca-Pérez, 2017). To explain this behavioral and ecological activity, different strategies are used by the anuran species to coexist in the same habitat: ontogenetically (Eterovick et al, 2010, de Menezes Gondim, 2013, foraging sites and competition inter and intraspecific (Duellman and Pyles, 1983;Toft, 1985), and by attracting females during the breeding season by strategic vocalization frequencies in males (Miyamoto, 1982).…”