One of the puzzles about the end-Cretaceous extinctions is why some organisms disappeared and others survived. A notable example is the differential extinction of ammonites and survival of nautilids, the two groups of co-occurring, externally shelled cephalopods at the end of the Cretaceous. To investigate the role of geographic distribution in explaining this outcome, we compiled a database of all the occurrences of ammonites and the nautilid genus Eutrephoceras in the last 0.5 m.y. of the Maastrichtian. We also included recently published data on ammonite genera that appear to have briefly survived into the Paleocene. Using two metrics to evaluate the geographic range of each genus (first, a convex hull encompassing all of the occurrences of each genus, and second, the maximum distance between occurrences for each genus), we documented that most ammonite genera at the end of the Maastrichtian were restricted in their geographic distribution, possibly making them more vulnerable to extinction. The geographic distribution of those genera that may have briefly survived into the Paleocene is significantly greater than that of non-surviving genera, implying that more broadly distributed genera were more resistant to extinction. This pattern is further emphasized by the broad distribution of Eutrephoceras, which matches that of the most widely distributed ammonites at the end of the Maastrichtian. However, even the most widely distributed ammonites eventually succumbed to extinction, whereas Eutrephoceras survived. Evidently, a broad geographic distribution may have initially protected some ammonites against extinction, but it did not guarantee their survival.
A 5.50 m thick interval of fossiliferous intensely bioturbated heterogenous glauconiferous sand of the upper Miocene Diest Formation is documented from a very large temporary outcrop just southeast of Antwerp International Airport (northern Belgium), allowing to observe lateral variations over several hundreds of meters and to collect many vertebrate and invertebrate fossils. This paper documents observations on lithology, sedimentary and post-sedimentary structures, and discusses the results of the multi-proxy analyses of the sediment (granulometry, glauconite content, clay mineralogy, Fe content and Fe3+/Fe2+ ratios), the interpretation of the trace fossil assemblage and the sedimentary structures as well as of the large-scale samplings of micro-, meso- and macrofossils. We evidence that the Diest Formation in the Antwerp area consists of two different lithological entities, and that this twofold character can be extrapolated to all previously recorded Deurne Member outcrops. A revised lithostratigraphic scheme for the Diest Formation in the Antwerp area is proposed, with the new Borsbeek member at the base and a redefined Deurne Member at the top.
Mesozoic marine reptiles went through a severe turnover near the end of the Triassic.Notably, an important extinction event affected ichthyosaurs, sweeping a large part of the group. This crisis is however obscured by an extremely poor fossil record and is regarded as protracted over the entire Norian-earliest Jurassic interval, for the lack of a more precise scenario. The iconic whale-sized shastasaurid ichthyosaurs are regarded as early victims of this turnover, disappearing by the middle Norian. Here we evaluate the pattern of this turnover among ichthyosaurs by analyzing the faunal record of two Rhaetian localities. One locality is Autun, eastern France; we re-discovered in this material the holotypes or partial 'type' series of
Rachitrema pellati, Actiosaurus gaudryi, Ichthyosaurus rheticus, Ichthyosaurus carinatus, andPlesiosaurus bibractensis; a revised taxonomic scheme is proposed. The second assemblage comes from a new locality: Cuers, southeastern France. Both these assemblages provide several lines of evidence for the presence of shastasaurid-like ichthyosaurs in the Rhaetian of Europe. These occurrences suggest that both the demise of shastasaurids and the sudden radiation of neoichthyosaurians occurred within a short time window; this turnover appear more abrupt but also more complex than previously postulated and adds a new facet of the endTriassic mass extinction.
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