The effects of anthropogenic disturbances on forest structure and plant diversity of secondary forest ecosystems were evaluated based on the classification by site factors in the montane regions of northeastern China. Forty-five sample plots containing 720 sub-plots of overstory species (8 m · 8 m), 1,440 quadrats of understory species (2 m · 2 m), and 1,440 quadrats of herbaceous layer species (1 m · 1 m) were clustered into five groups (G1, G2, G3, G4 and G5) by site variables with Hierarchical Cluster Analysis. Meanwhile, the disturbance levels corresponding to the five groups were determined according to the factors influencing human disturbances (D0-G2, D1-G1, D2-G3, D3-G5, and D4-G4). Species diversities of overstory, understory, herbaceous layer and overall species were evaluated using species number, Margalef index, Pielou index, Shannon-Wiener index, and Simpson index; and b-diversities (Whittaker index, Cody index). Basal area of stands exhibited a decreasing trend along the disturbance level due to a gradual increase in the extraction of timber by the human disturbances. The indices of species diversity suggested that overstory and understory species were distributed evenly among the groups or disturbance levels. There were no absolutely dominant tree species in the secondary forest ecosystems. The differences in site factors and the current disturbance intensities were not intense enough to lead to loose changes in overstory and understory species. The species diversity indices exhibited the maximum values at D3 (G5) for herbaceous layer species; this may suggest that the current disturbance intensities were intense enough to lead the changes of herbaceous layer species. Three rare and endangered species (Juglans mandshurica, Fraxinus mandshurica and Acanthopanax senticosus) were found within the secondary forests. These rare and endangered species appeared in each clustered group or disturbance level, which suggested that the current disturbance intensity in the study area was not strong enough to influence the distribution of rare and endangered species. The current disturbances in the secondary forests may not lead to a decrease in stability and complexity of the overstory and understory species, but the higher disturbance level may be intense enough to change the habitat fitness for the herbaceous species.

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Understory light is essential to the establishment and growth of understory plants and varies temporally and spatially within gaps. The previously defined gap light index (GLI) is a good model for assessing understory light levels, but it is time-consuming to determine gap coordinates, which are crucial to GLI, for numerous points within a gap. This paper introduces the geometric calculation (GeoCalc) of gap coordinates. GeoCalc quickly obtains gap coordinates for any specified point within a canopy gap and takes into account the tridimensional profile of the gap and the slope and aspect of the ground. The GeoCalc-based GLI was validated by the GLI derived from hemispherical photographs taken at 93 sampling points within seven natural gaps. Our results demonstrate that GeoCalc-based GLI was strongly positively correlated and not significantly differed from the GLI derived from hemispherical photographs. Next, to analyze gap light regimes and the effects of gap size, canopy height, and topography, three natural gaps of various size were selected and simulated as nine gaps with 1 and 1.5 times canopy height or on the opposite slope. Finally, we have summarized characteristics of GeoCalc-based GLI and its application.