prepared the samples for metabolomic analysis. R.V. and G.G. analyzed the metabolites in different mulberry cultivars and interpreted the data. N.T. and T.I. contributed to histochemical analysis and pulp preparation, respectively. M.Y. and N. prepared milled wood samples and M.Y. analyzed lignin structure by thioacidolysis. S.L., H.K., and J.R. prepared samples for NMR and performed NMR analysis and interpretation of the data. S.S. and N.M. analyzed the monomeric composition of cell wall polysaccharides. N and M.U. managed cultivated plants. S.K. wrote the