Five population‐specific response functions were developed from quadratic models for 110 populations of Pinus sylvestris growing at 47 planting sites in Eurasia and North America. The functions predict 13 year height from climate: degree‐days > 5 °C; mean annual temperature; degree‐days < 0 °C; summer‐winter temperature differential; and a moisture index, the ratio of degree‐days > 5 °C to mean annual precipitation. Validation of the response functions with two sets of independent data produced for all functions statistically significant simple correlations with coefficients as high as 0.81 between actual and predicted heights. The response functions described the widely different growth potentials typical of natural populations and demonstrated that these growth potentials have different climatic optima. Populations nonetheless tend to inhabit climates colder than their optima, with the disparity between the optimal and inhabited climates becoming greater as the climate becomes more severe. When driven by a global warming scenario of the Hadley Center, the functions described short‐term physiologic and long‐term evolutionary effects that were geographically complex. The short‐term effects should be negative in the warmest climates but strongly positive in the coldest. Long‐term effects eventually should ameliorate the negative short‐term impacts, enhance the positive, and in time, substantially increase productivity throughout most of the contemporary pine forests of Eurasia. Realizing the long‐term gains will require redistribution of genotypes across the landscape, a process that should take up to 13 generations and therefore many years.
Data points intensively sampling 46 North American biomes were used to predict the geographic distribution of biomes from climate variables using the Random Forests classification tree. Techniques were incorporated to accommodate a large number of classes and to predict the future occurrence of climates beyond the contemporary climatic range of the biomes. Errors of prediction from the statistical model averaged 3.7%, but for individual biomes, ranged from 0% to 21.5%. In validating the ability of the model to identify climates without analogs, 78% of 1528 locations outside North America and 81% of land area of the Caribbean Islands were predicted to have no analogs among the 46 biomes. Biome climates were projected into the future according to low and high greenhouse gas emission scenarios of three General Circulation Models for three periods, the decades surrounding 2030, 2060, and 2090. Prominent in the projections were (1) expansion of climates suitable for the tropical dry deciduous forests of Mexico, (2) expansion of climates typifying desertscrub biomes of western USA and northern Mexico, (3) stability of climates typifying the evergreen-deciduous forests of eastern USA, and (4) northward expansion of climates suited to temperate forests, Great Plains grasslands, and montane forests to the detriment of taiga and tundra climates. Maps indicating either poor agreement among projections or climates without contemporary analogs identify geographic areas where land management programs would be most equivocal. Concentrating efforts and resources where projections are more certain can assure land managers a greater likelihood of success.
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