Uranium-lead geochronology in detrital zircons and provenance analyses in eight boreholes and two surface stratigraphic sections in the northern Andes provide insight into the time of closure of the Central American Seaway. The timing of this closure has been correlated with Plio-Pleistocene global oceanographic, atmospheric, and biotic events. We found that a uniquely Panamanian Eocene detrital zircon fingerprint is pronounced in middle Miocene fluvial and shallow marine strata cropping out in the northern Andes but is absent in underlying lower Miocene and Oligocene strata. We contend that this fingerprint demonstrates a fluvial connection, and therefore the absence of an intervening seaway, between the Panama arc and South America in middle Miocene times; the Central American Seaway had vanished by that time.

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Tectonic collision between South America and Panama began at 23-25 Ma. The collision is signifi cant because it ultimately led to development of the Panamanian Isthmus, which in turn had wide-ranging oceanic, climatic, biologic, and tectonic implications. Within the Panama Canal Zone, volcanic activity transitioned from hydrous mantle-wedge−derived arc magmatism to localized extensional arc magmatism at 24 Ma, and overall marks a permanent change in arc evolution. We interpret the arc geochemical change to result from fracturing of the Panama block during initial collision with South America. Fracturing of the Panama block led to localized crustal extension, normal faulting, sedimentary basin formation, and extensional magmatism in the Canal Basin and Bocas del Toro. Synchronous with this change, both Panama and inboard South America experienced a broad episode of exhumation indicated by (U-Th)/He and fi ssion-track thermochronology coupled with changing geographic patterns of sedimentary deposition in the Colombian Eastern Cordillera and Llanos Basin. Such observations allow for construction of a new tectonic model of the South America-Panama collision, northern Andes uplift and Panama orocline formation. Finally, synchroneity of Panama arc chemical changes and linked uplift indicates that onset of collision and Isthmus formation began earlier than commonly assumed.

Closure of the Central American seaway was a local tectonic event with potentially global biotic and environmental repercussions. We report geochronological (six U/Pb LA‐ICP‐MS zircon ages) and geochemical (19 XRF and ICP‐MS analyses) data from the Isthmus of Panama that allow definition of a distinctive succession of plateau sequences to subduction‐related protoarc to arc volcaniclastic rocks intruded by Late Cretaceous to middle Eocene intermediate plutonic rocks (67.6 ± 1.4 Ma to 41.1 ± 0.7 Ma). Paleomagnetic analyses (24 sites, 192 cores) in this same belt reveal large counterclockwise vertical‐axis rotations (70.9° ± 6.7°), and moderate clockwise rotations (between 40° ± 4.1° and 56.2° ± 11.1°) on either side of an east‐west trending fault at the apex of the Isthmus (Rio Gatun Fault), consistent with Isthmus curvature. An Oligocene‐Miocene arc crosscuts the older, deformed and segmented arc sequences, and shows no significant vertical‐axis rotation or deformation. There are three main stages of deformation: 1) left‐lateral, strike‐slip offset of the arc (∼100 km), and counterclockwise vertical‐axis rotation of western arc segments between 38 and 28 Ma; 2) clockwise rotation of central arc segments between 28 and 25 Ma; and 3) orocline tightening after 25 Ma. When this reconstruction is placed in a global plate tectonic framework, and published exhumation data is added, the Central American seaway disappears at 15 Ma, suggesting that by the time of northern hemisphere glaciation, deep‐water circulation had long been severed in Central America.